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The yawn reflex has long been observed to be contagious. In 1508, Erasmus wrote, "One man's yawning makes another yawn., Often, if one person yawns, this may cause another person to "empathetically" yawn. Observing another person's yawning face (especially his/her eyes), even reading, or thinking about yawning, or looking at a yawning picture can cause a person to yawn. The proximate cause for contagious yawning may lie with mirror neurons in the frontal cortex of certain vertebrates, which, upon being exposed to a stimulus from conspecific (same species) and occasionally interspecific organisms, activates the same regions in the brain. Mirror neurons have been proposed as a driving force for imitation which lies at the root of much human learning such as language acquisition. Yawning may be an offshoot of the same imitative impulse.
A 2007 study found that young children with autism spectrum disorders do not increase their yawning frequency after seeing videos of other people yawning, in contrast to typically developing children. In fact, the autistic children actually yawned less during the videos of yawning than during the control videos. This supports the claim that contagious yawning is related to empathic capacity.

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The relationship between yawn contagion and empathy is strongly supported by a 2011 behavioral study, conducted by Ivan Norscia and Elisabetta Palagi (University of Pisa, Italy). The study revealed that - among other variables such as nationality, gender, and sensory modality - only social bonding predicted the occurrence, frequency, and latency of yawn contagion.[29] As with other measures of empathy, the rate of contagion was found to be greatest in response to kin, then friends, then acquaintances, and lastly strangers.[29] Related individuals (r≥0.25) showed the greatest contagion, in terms of both occurrence of yawning and frequency of yawns.[29] Strangers and acquaintances showed a longer delay in the yawn response (latency) compared to friends and kin.[29] Hence, yawn contagion appears to be primarily driven by the emotional closeness between individuals.[29]
Yawning has been observed among various primates. In these cases the yawn is a threat gesture, a way of maintaining order in the primates' social structure. Specific studies were conducted on chimpanzees[30] and stumptail macaques.[31] A group of these animals was shown a video of other conspecifics yawning; both species yawned as well. This helps to partly confirm a yawn's "contagiousness".
The Discovery Channel's show Mythbusters also tested this concept. In their small-scale, informal study they concluded that yawning is contagious.[32]
Gordon Gallup, who hypothesizes that yawning may be a means of keeping the brain cool, also hypothesizes that "contagious" yawning may be a survival instinct inherited from our evolutionary past. "During human evolutionary history, when we were subject to predation and attacks by other groups, if everybody yawns in response to seeing someone yawn the whole group becomes much more vigilant and much better at being able to detect danger."[17]
A study by the University of London has suggested that the "contagiousness" of yawns by a human will pass to dogs. The study observed that 21 of 29 dogs yawned when a stranger yawned in front of them, but did not yawn when the stranger only opened his mouth.[4] A recent study from Lund University showed that dogs, like humans,[33] develop a susceptibility to contagious yawning gradually, and that while dogs above 7 months 'catch' yawns from humans, younger dogs are immune to contagion.[34] The study also indicated that nearly half of the dogs responded to the human's yawn by becoming relaxed and sleepy, suggesting that the dogs copied not just the yawn, but also the physical state that yawns typically reflect

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